P-322's intake is on top of the boom ahead of the wing leading edge. Air comes in and make a 90* down into the carb. All the intake plumbing for the turbo is simply left out; the outer leading edge can probably be simplified aka made lighter is it isn't being used as an intercooler (or turned into fuel tank to extend range).
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P-322 to Lightning I is the same change as P-40B to D: i.e. change from Allison C to F engines. The problem with the "C" nose is it uses: a: an internally toothed gear for reduction box (needs custom machining equipment) and a huge plain bearing. With number of engines being ordered and power increases to the engine, it was more challenging to increase capacity than to change to the 2 externally toothed gears. So, Allison changed production to the F nose*. But the actual thrust line doesn't change that much (Curtis/Lockheed still probably need to manufacture new cowls).
Could the low supercharger ratios be part of the contract for the French i.e. They wanted engines to be able to run on 87 (or worse) fuel. Then Allison was just fulfilling the contract they had in hand. I know there is a lot of bad blood between Lockheed and British over what was ordered versus what was needed.
*It makes commonality with the Allison E series easier too. C nose engines drove the supercharger via a shaft from the reduction gear. But the P-39 doesn't have a reduction gear on the engine, adding the reduction gears - crankshaft and internally tooth gear to drive the supercharger are expensive solution. Taking the drive off the back end of the crankshaft for the supercharger is a common solution.
The ~200 Merlins lost in Summer '40 when RR workers went on strike in middle of BoB would have powered 100+ Lightning Is... (You can see the dip in numbers
G
Geoffrey Sinclair provides). But there was no better time to get concessions from gov't for better housing, etc.