Anyone know the story of this B-29?

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Not sure about the XB-54 test bit.

The aircraft is 42-24441 modified under Projetc S68 and had those two Emerson Model 136 jowl barbettes each with a 0.5" MG. But it also had the GE remote controlled turrets replaced with manned Martin turrets in the upper positions and manned Sperry A-2 ball turret in the lower forward position and a similar modified A-13 ball turret in the lower aft position. The waist blisters were replaced with manned flexible 0.5" MG. The above photo is one of a series from the Boeing archives dated to Aug - Oct 1944. See Detail & Scale Vol 10 B-29 Superfortress Part 1.
 

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