The Hornet prototype RR 915 first flew on 28 July 1944 with Geoffrey de Havilland Jr. at the controls. Powered by twin Merlin engines, it was the fastest piston-engined fighter in Royal Air Force service. The Hornet also has the distinction of being the fastest wooden aircraft ever built and the second fastest operational twin propeller-driven aircraft — being slightly slower than the unconventional German Dornier Do 335 of 1945.
The prototype achieved 485 mph (780 km/h) in level flight, which came down to 472 mph (760 km/h) in production aircraft.
The first 10 Do 335 A-0s were delivered for testing in May. By late 1944, the Do 335 A-1 was on the production line. This was similar to the A-0 but with the uprated DB 603 E-1 engines and two underwing hardpoints for additional bombs, drop tanks or guns. It was capable of a maximum speed of 763 km/h (474 mph) at 6,500 m (21,300 ft) with MW 50 boost, or 686 km/h (426 mph) without boost, and able to climb to 8,000 m (26,250 ft) in under 15 minutes. Even with one engine out, it could reach about 563 km/h (350 mph).