World War II fighter armament: what was too light, what was overkill, what was the Goldilocks zone?

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From the Wikipedia entry for the Gloster Gladiator, 4 x.303 MG's were able to kill single and multi engine, metal construction, radial engine aircraft from Italy, Japan , and the Soviet Union. It looks as though having all 4 functioning at any time was quite unusual.
 
I think the increase to 8 was based on the spread pattern used and a two second burst of fire.
 

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