The P-39 was a slick aircraft (low drag) but it was heavy. It did do pretty well in miles per gallon but a lot of these aircraft were pretty close together and does a 10-20% change in range really amount to all that much operationally?
Like if plane "A" has a operational radius if 100 miles and plane "B" has an operational radius of 120 miles and you are dealing with the English channel which can be 100 miles wide (or more) in some spots what is the practical difference?
Even on the Eastern front, sometimes a battle will be that grey zone where plane B can reach it from a certain airfield but plane A can't but how often is that going to happen where there isn't one or more airfields closer to the battle that Plane A couldn't stage through, either going or coming.
Here is the range chart for a P-39Q-1.
http://zenoswarbirdvideos.com/Images/P-39/P39FOIC.pdf
down low (0-6000ft) at 250-260mph it can do about 260 miles after subtracting 6-17 gallons for warm up and take off. WIth 120 gallon tanks (105 usable) it should do about 390 miles at that speed but there is no allowance for either combat or for finding home base and landing. Slower speeds give more range and max continuous less.
In British terms the P-39Q comes out like this for 5 minute fuel usage.
Cruise condition..................5 min at Military power...........5 min at WEP
Max cruise 3.125mpg.................36 miles................................44 miles
Intermed (2nd column)...............44 miles................................53 miles
most economical........................74 miles................................90 miles
Fractions rounded off.
Spitfires were all over the place in regards to fuel capacity. with 102 US gallons on the early ones. 146 US gallons on the MK VII & VIII, 114 US gallons on some late MK IXs
And then you have the fact that the Spitfire
could operate with the 36 gallon Slipper tank (perform all combat maneuvers?) , how often this was done I don't know.
Apparently it was only the Griffon engined MK XIV that used the 108 US gallon slipper tank in combat?
Unknown to me is if the 54 US gallon slipper tank was ever used in combat.
By used in combat I mean the plane did not drop it when engaging the enemy. I have no idea if there were fuel feed problems with any of theses slipper tanks or if there was a problem with fuel sloshing around if the tank was part full. (how much internal baffling there was? Or how strong the attachment points were.
My Apologies to our commonwealth members but I figured I would keep all fuel in US gallons to help eliminate confusion.