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Wings had the SPACE to mount fuel tanks where the wing guns were mounted. "
You would have space in two dimensions. Now the question is it was smart.
Fuel tanks fitted between the forward spar and the Main spar, the distance narrows between the spars the further out you go.
The wing gets thinner there further out you go. The wing at the W-2 position indicated on the drawing is less than 8 inches thick at max, it is a bit thicker where the machine guns are.
Self sealing fuel tank material is pretty much constant thickness and weight per sq ft. A long, skinny, thin tank is going to use a lot more self sealing material than a short fat tank of the same capacity. a tank with an assumed thickness of 6.5 inches (allowing for Self sealing material as a guess, corrections welcome) you need almost 2 sq feet of tank to hold
ONE gallon.
Granted that is worst case, but it shows that fuel tanks in the last few feet of a wing are poor proposition. Lots of tank weight for not that big an increase in fuel. A tank at mid point in the wing may hold 50 % more fuel per sq ft of wing area than tank in the outer wing for the same weight of self sealing material due to being thicker.
when outer wing tanks are full role response is poor.
The post-war P-39 racing planes held 100gal in the outer wings (50gal per side). I'm saying that 30 additional gallons would be plenty (150gal total) which would be only 15gal per side. The entire area where the tanks would go in the outer wing is clear area, not broken up by wing ribs as in the normal P-39 wing tanks. Roll would be no worse than with wing guns and full ammo boxes, weight would be about the same.
"N was not a low altitude plane."
Engine peaked at 15,000ft or so. Yes there were lower altitude planes but many fighters had engines that peaked in the 20,000ft foot range.
Spitfire II engine peaked 17,500ft in the summer of 1940.
I suggest you should not worry so much about the specific performance of each engine, but more about the performance of the plane that the engine was installed in. P-39 had less drag than Spitfire.