At least someone understands the argument, MkV's used slippers over France that didn't increase their range but instead the extra fuel allowed for a higher cruising speed so it made them harder to bounce, another advantage of extra fuel.We can safely assume that the 29 gals from the rear tank is perhaps 10-15 gals before the German (or other Axis) A/C; or empty, since the manual prescribes that the rear tank is to be used up 1st. Once the drop tank is jettisoned, the influence at the CoG coming from the rear tank is negligible. Some re-shuffling of the counterweights might've helped, too.
Th early (talk 1941/42) and regular use of 29 gal tank on the Spitfires also puts the Spitfire IX and VIII in a better position, range- and usability-wise.
Spitfires with more fuel onboard can cruise on faster speed where the danger is suspected, so the German fighters have the harder time to dictate terms of combat.