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Hope the ball turret-gunner was seated next to the engineer! ;)
As a general rule, assembly ships had only a limited crew for flying and navigation, hence no gunner as the ship returned to base after formation completed..
However, as exception to the rule, according to Wiki, there were instances when Spotted Ass Ape flew to Germany with the 458th formation.
 
I take it those are lights or strobes or flare dischargers to assist other pilots in locating the aircraft in the middle of the I
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