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That's right, the lead plane was based on (ie. rebuilt with parts mainly from) B-17D 40-3095, one of those based in the Philippines at the beginning of the war. One of the others was based on B-17E 41-2471, incompletely destroyed by burning when abandoned on Java in late Feb. 1942. The other one's identity is unknown, it apparently only appears in that picture. A Japanese book on captured Allied a/c doesn't mention it. It's conceivable that's a retouched photo, but in any case both E's in that picture are early ones (like 41-2471) with Bendix remote control belly turrets, rather than Sperry ball turrets.The lead B-17 is a "D" model and the ones following are "E" or "F" models.
Here a picture that really confusing me. Just where is this pickture taken?